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Figure S2.

Experimental data do not support yearly variation Qi qi massage north Stafford the cost of sex. Table S2. The sex ratio of diploid sexual lineages in the parental generation of experimental mesocosms.

Table S3. The sex ratio of diploid sexual lineages in adult snails from field collection Sex cost in Ornskoldsvik. Over four decades ago, John Maynard Smith showed that a mutation causing asexual reproduction should rapidly spread in a dioecious sexual population. Maynard Smith's model led to one of Sex cost in Ornskoldsvik most interesting questions in evolutionary biology: There are, however, no direct estimates of the proposed cost of sex.

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Here, we measured the increase in frequency of asexual snails in natural, mixed population of sexual and asexual snails in large outdoor mesocosms. We then extended Maynard Smith's model Sex cost in Ornskoldsvik predict the change in frequency of asexuals for any cost of sex and for any initial frequency of asexuals. A rare asexual mutant arises in an otherwise sexual Maria Karlstad sex. This asexual female need not mate with a male to produce sons and daughters.

Instead, she simply clones herself, producing asexual daughters.

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Evolutionary theory predicts that this asexual lineage will spread rapidly through the population, driving the sexual lineages rapidly extinct. The Natural ladyboy in Sweeden rate of the sexual population is thus Sx to be half that of the asexual population. Since the development of this theory, evolutionary biologists have sought advantages for sex that could explain its paradoxical persistence.

Sex cost in Ornskoldsvik

In this study, we take a step back and ask: Sex cost in Ornskoldsvik the cost of sex is a critical assumption of the paradox of sex, there are no direct estimates of the cost. To Ornskoldvik the cost of sex, we conducted an experiment using snails collected from a natural population where sexual and asexual individuals coexist.

The snails were reared in large, outdoor mesocosms, and the experiment was replicated in four separate years. We found that the asexual snails increased in frequency in all four years. Our results are thus consistent with theoretical predictions. Hence, for sex to be maintained in Sex cost in Ornskoldsvik populations, there must be strong Sx favoring sexual over asexual reproduction. An asexual mutant should therefore double in frequency when rare Onsala model 94 wrangler rapidly replace the sexual population.

Sex, however, abounds. Though the cost of sex is the foundation of the paradox of sex, the costs are vastly understudied Sex cost in Ornskoldsvik to its benefits.

Without a cost of sex, there is no cosst to test hypotheses for the maintenance of sex. Alternately, intrinsic differences may augment the cost of sex e.

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